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However, MGST1 was observed to decrease in middle- and old-age rat liver compared to young rats, consistent with the upregulation of miR-34a and miR-93 during aging and downregulation of their targets SP1 and Nrf2 which are the transcription factors for MGST1 (Li, Muthusamy, Liang, Sarojini, & Wang, 2011)